Costa Rica

Male slaty-tailed trogon (Trogon massena), Kelady, Costa Rica

Tiny terrestrial crab (sp.), Las Nubes, Costa Rica

Bay-headed Tanager, Santa Elena, Costa Rica (Tangara gyrola) CR SP - Tangara Cabecicastana

Tree fern, one of the oldest plants on earth and foundation for the Earth's first forests, Las Nubes, Costa Rica

Rainforest tree backlight at sunrise in the Las Nubes Reserve, Costa Rica

Jewel-like raindrops on (sp.) flower (wild, not cultivated), Las Nubes, Costa Rica

A driver ferries sugar cane to the mill in Santa Elena, Costa Rica.

The dramatic muddy brown waters of the Rio Susio (Dirty River) flowing through Braulio Carillo National Park, Costa Rica.

Quizarra resident Cesar Ivan Rojas Arias handles a (sp.) bird that will be banded as part of a York University research project, Las Nubes, Costa Rica

A quality-control worker cradles a handful of ultra-premium coffee beans at the CoopeAgri factory in San Isidro, Costa Rica. These are as good as it gets - large, red, ripe, organic, shade-grown, high-elevation beans - and command a high price.

Bumble bee grasshopper, CoopeSilencio, Costa Rica

Forest abstract with a leaf mimicing an animal form, taken in a rain storm, Las Nubes, Costa Rica

Rufus-collared sparrow (Zonotrichia capensis) CR SP – Chingolo, on the summit of Volcan Poas, Costa Rica

Caterpillar that has been taken over by a parasitic wasp, Las Nubes, Costa Rica

Macro view of a butterfly (sp.) along a small creek in the Las Nubes Reserve, Costa Rica.

Macro view of a small spider (sp.) with a spectacularly decorated abdomen. In the Las Nubes Reserve, Costa Rica.

Male summer tanager (Pirangra ruba), CR SP Tangara Veranera, San Isidro, Costa Rica.

Large backlit ferns in primary forest, Las Nubes, Costa Rica

View of forested slope in the Penas Blancas valley from the Las Nubes Reserve, Costa Rica

Detail of tree fern fronds, Las Nubes, Costa Rica

Dramatic late afternoon clouds over the Valle General, Las Nubes, Costa Rica

The photographer's assistant poses in a tributary to the Rio Penas Blancas, Las Nubes, Costa Rica

A group of Costa Ricans riding horses along the beach in Marino Ballena National Park, Costa Rica.

Large backlit rainforest leaf, Las Nubes, Costa Rica

The sun shines through dramatic stormclouds onto the Pacific Ocean. Near Quepos, Costa Rica

Eye to eye with grazing cows, with the rain forests of the Las Nubes Reserve in the background. Near San Isidro, Costa Rica.

The bible of Costa Rican birdwatching - "A Guide to the Birds of Costa Rica" - co-authored by the late Dr. Alexander Skutch, bearing his signature and that of the artist Dana Gardner. Los Cusingos, Quizarra, Costa Rica.

Fog envelopes the rainforest, Las Nubes, Costa Rica

High water level and sediment load in the Rio Penas Blancas after a torrential rain, Las Nubes, Costa Rica

Large, beautifully patterned butterfly (sp.) camoflagued on the underside of a large fallen tree trunk, Las Nubes, Costa Rica

Dramatic late afternoon clouds over the Alexander Skutch Biological Corridor, near San Isidro, Costa Rica.

Large bromeliad showing lotus pattern of leaves, Las Nubes, Costa Rica

Elegant symmetry in a plant on the rainforest floor, Las Nubes, Costa Rica

Male red-legged honeycreeper (Cyanerpes cyaneus), CR SP Mielero Patirrojo, La Escondita, Quizarra, Costa Rica

Mono Cariblanco (Cebus Capucinus) in the Los Cusingos Reserve, Costa Rica

Exquisitely detailed grasshopper (sp.), Las Nubes, Costa Rica

Detail of the Rio Penas Blancas, Las Nubes, Costa Rica

Baby reptile (sp.), 6 cm with tail, Rio Penas Blancas, Skutch Corridor, Costa Rica

Salamander (sp.) found in an unusual position on a dry tree trunk in bright sunlight, Las Nubes, Costa Rica

Clouds roll in over primary forest, Las Nubes, Costa Rica

Vistas de Chirripo, the closest housing development to the Las Nubes Reserve, high-priced and owned by foreigners, Las Nubes, Costa Rica

Dramatic late afternoon clouds over the Valle General, Las Nubes, Costa Rica

Backlit cacao leaves, the plant used to make chocolate, on the Los Cusingos Reserve, Costa Rica.

Lichen (usnia?) blankets a rainforest tree, Las Nubes, Costa Rica

Local residents pose on their horse, Santa Elena, Skutch Corridor, Costa Rica

Dainty flowers of the (sp.) plant, Las Nubes, Costa Rica

(Non-breeding) Laughing gulls (Larus atricilla), CR SPGaviota Reidora, Marino Ballena National Park, Costa Rica

Dramatic yellow light at dusk over classic view of the Skutch Corridor, Costa Rica

Sugar cane workers from Nicaragua, Quizzara, Costa Rica

White-ruffed Manakin (Corapipo leucorrhoa), sp. Saltarin Gorgiblanco. Las Nubesa Reserve, Costa Rica

Medium-sized reptile (12cm), Los Cusingos Sanctuary, Costa Rica

Blue-gray tanager (Thraupis episcopus), CR SP Viuda, Tangara Azuleja, Los Cusingos Reserve, Costa Rica

Distant view of the Las Nubes Reserve, Costa Rica

Unidentified flowers (sp.) in regenerating forest, Las Nubes, Costa Rica

Black vultures (Coragyps atratus) in a large palm tree on the beach south of Puerto Jimenez, Osa Peninsula, Costa Rica

Rainforest at the edge of the Las Nubes Reserve, Costa Rica.

Macro view of sap dripping from a branch on a palm tree in the cloud forests of Volcan Poaz, Costa Rica.

Daddy long legs (sp.), Las Nubes, Costa Rica

Man's two best friends, Quizarra, Costa Rica

Speckled tanager (Tangara guttata), CR SP Zebra, Tanagra Moteada, Los Cusingos Reserve, Costa Rica

Strange translucent gel on a tree branch, with what appear to be seeds suspended inside. It came from a source above. Las Nubes, Costa Rica

Typical view of countryside in the Skutch Corridor, Costa Rica

Early morning light on partially deforested lands in the Alexander Skutch Biological Corridor, near the Las Nubes Reserve, Costa Rica.

Large frog (sp.), Los Cusingos Sanctuary, Costa Rica

A group of Costa Ricans riding horses along the beach in Marino Ballena National Park, Costa Rica.

Bromeliad on the summit of Volcan Irazu, the highest volcano in Costa Rica.

One section of an enormous pineapple plantation near San Isidro, Costa Rica. This large industrial operation is heavily dependant on chemicals and the plantation is a complete monoculture.

Silhouetted moth (sp.) atop a leaf as seen from underneath, Las Nubes, Costa Rica

A spiky, histamine-laden moth caterpillar, likely Hylesia continua, painful to the touch. Los Cusingos Reserve, Costa Rica.

Boa constrictor (Boa constrictor), SP Becquer, Los Cusingos Sanctuary, Costa Rica

Rainbow over the Alexander Skutch Biological Corridor, Costa Rica.

Unusual beetle, in its playing dead phase. Fine detail of underside is visible at full-size. Finca David, Quizarra, Costa Rica

Abstraction of palm fronds, Skutch Corridor, Costa Rica